Buster (Oscar® winner Matthew McConaughey) has turned the New Moon Theater into a local hit, but Buster has his eyes on a bigger prize: Debuting a new show at the Crystal Tower Theater in glamorous Redshore City.

But with no connections, Buster and his cast—including harried mother pig Rosita (Oscar® winner Reese Witherspoon), rocker porcupine Ash (Scarlett Johansson), earnest gorilla Johnny (Taron Egerton), shy elephant Meena (Tori Kelly) and, of course, porcine provocateur Gunter (Nick Kroll)—have to sneak their way into the world-famous Crystal Entertainment offices, run by ruthless mogul wolf Jimmy Crystal (Emmy winner Bobby Cannavale).

In a desperate attempt to get Mr. Crystal’s attention, Gunter spontaneously pitches an outrageous idea that Buster quickly runs with, promising that their new show will star lion rock legend Clay Calloway (Bono). Problem is, Buster has never met Clay, an artist who shut himself away from the world more than a decade ago after the loss of his wife. Even worse, Buster has failed to realize that Mr. Crystal is an egocentric gangster who would rather drop somebody off the roof of a building than be lied to.

As Gunter helps Buster dream up an out-of-this-world theatrical masterpiece, and the pressure (and sinister threats) from Mr. Crystal mount, Rosita’s leading role in the show gets ripped away and handed to Mr. Crystal’s spoiled daughter, Porsha, played by Grammy-nominated artist Halsey. Desperate to save the show, and his life, Buster embarks on a quest to find Clay and persuade him to return to the stage.

What begins as a dream of big-time success becomes an emotional reminder of the power of music to heal even the most broken heart.